substrate substratethe surface or medium on which an organism lives or grows.Related Articles: Glossary: essential fish habitat (EFH)How do marine invertebrates produce sounds?Sea Urchin (Kina)Marine invertebrate communicationMedia BackgrounderHow do marine invertebrates detect sounds?Tutorial Can Animals Sense These Sounds Part IIWind TurbineTutorial: Wind TurbinePile Driving Chris Knowlton2017-02-26T20:10:08-05:00