substrate substratethe surface or medium on which an organism lives or grows, or the material on the bottom of the ocean. Related Articles: Glossary: essential fish habitat (EFH)Underwater Sound and Coral Reef Restoration – September 14, 2022Considerations for “Contained” Studies in Laboratory or Captive SettingsWhat lies beneath: How benthic communities affect acoustic propagation – April 29, 2026What components of sound are used for hearing?How do marine invertebrates produce sounds?Sea Urchin (Kina)Potential effects of sound on marine invertebratesMarine invertebrate communicationPile driving Chris Knowlton2019-09-10T09:41:25-05:00