substrate substratethe surface or medium on which an organism lives or grows, or the material on the bottom of the ocean. Related Articles: Glossary: essential fish habitat (EFH)Underwater Sound and Coral Reef RestorationConsiderations for “Contained” Studies in Laboratory or Captive SettingsWhat components of sound are used for hearing?How do marine invertebrates produce sounds?Sea Urchin (Kina)Potential effects of sound on marine invertebratesMarine invertebrate communicationPile drivingMedia Backgrounder Chris Knowlton2019-09-10T09:41:25-05:00